Haplogroup O-M175

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Haplogroup O-M175
File:Haplogrupo O (ADN-Y).PNG
Possible time of origin41,750 (95% CI 30,597<-> 46,041) years ago[1]

44,700 or 38,300 ybp[2]
Coalescence age33,943 (95% CI 25,124 <-> 37,631) years (Karmin 2022[1])

35,000 or 30,000 years ago depending on mutation rate[2]
Possible place of originSoutheast Asia or East Asia
AncestorNO
DescendantsPrimary:
O1 (O-F265); O2 (O-M122) 
Secondary:
O1a (O-M119); O1b (O-M268);
O2a (O-M324); O2b (O-F742)
Defining mutationsM175 (+ numerous other SNPs).[3]

Haplogroup O, also known as O-M175, is a human Y-chromosome DNA haplogroup. It is primarily found among populations in Southeast Asia and East Asia. It also is found in various percentages of populations of the Russian Far East, South Asia, Central Asia, Caucasus, Crimea, Ukraine, Iran, Oceania, Madagascar and the Comoros. Haplogroup O is a primary descendant of haplogroup NO-M214. The O-M175 haplogroup is very common amongst males from East and Southeast Asia. It has two primary branches: O1 (O-F265) and O2 (O-M122). O1 is found at high frequencies amongst males native to Southeast Asia, Taiwan, the Japanese Archipelago, the Korean Peninsula, Madagascar and some populations in southern China and Austroasiatic speakers of India. O2 is found at high levels amongst Han Chinese, Tibeto-Burman populations (including many of those in Yunnan, Tibet, Burma, Northeast India, and Nepal), Manchu, Mongols (especially those who are citizens of the PRC), Koreans, Vietnamese, Filipinos, Japanese, Thais, Polynesians, Miao people, Hmong, the Naiman tribe of Kazakhs in Kazakhstan,[4] Kazakhs in the southeast of Altai Republic,[5] and Kazakhs in the Ili area of Xinjiang.[6]

Origins

Haplogroup O-M175 is a descendant haplogroup of Haplogroup NO-M214, and first appeared according to different theories either in Southeast Asia (see Rootsi 2006, TMC 1998, Shi 2005, and Bradshaw Foundation) or East Asia (see ISOGG 2012) approximately 40,000 years ago (or between 31,294 and 51,202 years ago according to Karmin et al. 2015).[7][8] Haplogroup O-M175 is one of NO-M214's two branches. The other is Haplogroup N, which is common throughout North Eurasia.

Distribution

This haplogroup appears in high to moderate frequencies in most populations in both East Asia and Southeast Asia, and it is almost exclusive to that region. It is almost nonexistent in Western Siberia, Western Asia, Europe, most of Africa, India and the Americas, where its presence may be the result of recent migrations. However, certain O subclades do achieve significant frequencies among some populations of Central Asia, South Asia, and Oceania. For example, one study found it at a rate of 67% among the Naimans, a tribe in Kazakhstan,[4] even though the rate among Kazakhs in general is only about 3.3% to 10.8%(Wells et al. 2001).[4][9] (It is notable that 75% of cases of haplogroup O-M175 observed in the Kazakh sample of Ashirbekov et al. 2017, of which 10.8% have been found to belong to haplogroup O-M175, have been contributed by the Naimans themselves; only 3.1% of the remainder of the Kazakh sample with the Naimans excluded belong to haplogroup O-M175.) It has been estimated that 25% of the entire male population of the world carries different subclades of O.[8][10] Karafet et al. (2015) have assigned the Y-DNA of 46.2% (12/26) of a sample of Papuan from Pantar Island to haplogroup NO-M214;[11] considering their location in the Malay Archipelago, all or most of these individuals should belong to haplogroup O-M175. An association with the spread of Austronesian languages in late antiquity is suggested by significant levels of O-M175 among island populations of the South Pacific and Indian Ocean, including the East African littoral. For example, Haplogroup O-M50 has even been found in Bantu-speaking populations of the Comoros along 6% of O-MSY2.2(xM50),[12] while both O-M50 and O-M95(xM88) occur commonly among the Malagasy people of Madagascar with a combined frequency of 34%.[13][14] O-M175 has been found in 28.1% of Solomon Islanders from Melanesia.[15] 12% of Uyghurs (Wells et al. 2001), 6.8% of Kalmyks[16] (17.1% of Khoshuud, 6.1% of Dörwöd, 3.3% of Torguud, 0% of Buzawa), 6.2% of Altaians (Kharkov et al. 2007), 4.1% of Uzbeks on average but Uzbeks from Bukhara 12.1%, Karakalpaks (Uzbekistan) 11.4%, Sinte (Uzbekistans) 6.7% (Wells et al. 2001) and 4.0% of Buryats.[17] In the Caucasus region it has been found in the Nogais 6%[18] but 5.3% in the Karan Nogais, it is also found in the Dargins of Dargwa speakers at 2.9%.[19] In the Iranic population, it is found in Iranian (Esfahan) at 6.3% (Wells et al. 2001), 8.9% of Tajiks in Afghanistan[20] 4.2% in the Pathans in Pakistan (Firasat 2007) but 1% in Afghanistan,[citation needed] 3.1% in Burusho (Firasat 2007). Haplogroup O-M175 ranges in various moderate to high frequencies in the ethnic minorities of South Africa. The frequency of this haplogroup is 6.14% in the Cape colored population,[21] 18% in Cape Coloured Muslim, 38% in Cape Indian Muslims and 10% in Cape Other Muslims.[21] It's found 11.5% in the Réunion Creole.[22] Haplogroup O-M175 had also been found in Latin America and Caribbean as a result of massive Chinese male migration from the 19th century. It was found in the Jamaicans at 3.8%,[23] and in Cubans, 1.5%.[24] Haplogroup O-M175 has been found in 88.7% of Asian American. 1.6% in Hispanic American, White Americans 0.5%, and 0.3% in African American.[25] Another study gives 0.5% African American.[26] Among the sub-branches of haplogroup O-M175 are O-M119(O1a), O-M268(O1b), and O-M122(O2). Y Haplogroup O3-M122 makes up the majority of Jadoon's males, the same haplogroup carried by the majority (50-60%) of Han Chinese. 82.5% of Jadoon men carrying Q-MEH2 and O3-M122 which are both of East Asian origin. O3-M122 was absent in the Sayyid (Syed) population and appeared in low numbers among Tanolis, Gujars and Yousafzais. There appears to be founder affect in the O3-M122 among the Jadoon.[27][28][29] 76.32% of Jadoon men carry O3-M122 while 0.75% of Tanolis, 0.81% of Gujars and 2.82% of Yousafzais carry O3-M122.[30][31] Russians in China East Asian haplogroup O made up 58% of their Y haplogroup. O3-M122 specifically made up 47% of the Russian sample.[32] The East Asian Y haplogroup O3-M122 was found in 47% of Russian males in China. In another test the East Asian paternal Y Haplogroup O made up 58% of Russian males samples in China.[33] Haplogroup O was found in 1%-1.2% of Persians in one sample.[34][35][36][better source needed][37][better source needed] O3-M122 is the commonly shared genetic signature of Sino-Tibetan speaking ethnicities.[38]

O-M175*

A broad survey of Y-chromosome variation among populations of central Eurasia found haplogroup O-M175(xM119,M95,M122) in 31% (14/45) of a sample of Koreans and in smaller percentages of Crimean Tatars (1/22 = 4.5%), Tajiks (1/16 = 6.25% Dushanbe, 1/40 = 2.5% Samarkand), Uyghurs (2/41 = 4.9%), Uzbeks (1/68 = 1.5% Surxondaryo, 1/70 = 1.4% Xorazm), and Kazakhs (1/54 = 1.9%) (Wells et al. 2001). However, nearly all of the purported Korean O-M175(xM119,M95,M122) Y-chromosomes may belong to Haplogroup O-M176,[Note 1] and later studies do not support the finding of O-M175* among similar population samples (Xue 2006, Kim 2011). The reported examples of O-M175(xM119,M95,M122) Y-chromosomes that have been found among these populations might therefore belong to Haplogroup O-M268*(xM95,M176) or Haplogroup O-M176 (O1b2). A study published in 2013 found O-M175(xM119, M95, M176, M122) Y-DNA in 5.5% (1/18) Iranians from Teheran, 5.4% (2/37) Tajiks from Badakhshan Province of Afghanistan, and 1/97 Mongols from northwest Mongolia, while finding O-M176 only in 1/20 Mongols from northeast Mongolia.[39]

O-F265 (O1)

O1a-M119 and O1b-M268 share a common ancestor, O1-F265 (a.k.a. O-F75) approximately 33,181 (95% CI 24,461 to 36,879) YBP.[1][40] O1-F265, in turn, coalesces to a common ancestor with O2-M122 approximately 33,943 (95% CI 25,124 to 37,631) YBP.[1] Thus, O1-F265 should have existed as a single haplogroup parallel to O2-M122 for a duration of approximately 762 years (or anywhere from 0 to 13,170 years considering the 95% CIs and assuming that the phylogeny is correct) before breaking up into its two extant descendant haplogroups, O1-MSY2.2 and O1b-M268.

O-M119 (O1a)

O-M119 (which was known briefly as O-MSY2.2, until the SNP MSY2.2 was found to be unreliable) is found frequently in Austronesian-speaking people, with a moderate distribution in southern and eastern Chinese and Kra-dai peoples.

O-M268 (O1b)

O-M122 (O2)

Found frequently among populations of East Asia, Southeast Asia, and culturally Austronesian regions of Oceania, with a moderate distribution in Central Asia (Shi 2005).

O-M324 (O2a)
O-F742 (O2b)

Language families and genes

Haplogroup O is associated with populations which speak Austric languages. The following is a phylogenetic tree of language families and their corresponding SNP markers, or haplogroups, sourced mainly from Edmondson 2007 and Shi 2005. This has been called the "Father Tongue Hypothesis" by George van Driem (van Driem 2011). It does not appear to account for O-M176, which is found among Japanese, Korean, and Manchurian males.

(M175) 
O2 (M122) 
 Sino-Tibetan O2a2b1 (M134) 

 Sinitic O2a2b1a2 (F114) 

 Sino-Tibetan O2a2b1a1 (M117) 

 Hmong–Mien O2a2a1a2 (M7) 

 Hmong (Miao) 

 She (Ratliff 1998

 Mien (Yao) 

O1 (F265) ("Austric") 

Phylogenetics

Phylogenetic history

Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium (YCC). They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being, above all, timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.

YCC 2002/2008 (Shorthand) (α) (β) (γ) (δ) (ε) (ζ) (η) YCC 2002 (Longhand) YCC 2005 (Longhand) YCC 2008 (Longhand) YCC 2010r (Longhand) ISOGG 2006 ISOGG 2007 ISOGG 2008 ISOGG 2009 ISOGG 2010 ISOGG 2011 ISOGG 2012
O-M175 26 VII 1U 28 Eu16 H9 I O* O O O O O O O O O O
O-M119 26 VII 1U 32 Eu16 H9 H O1* O1a O1a O1a O1a O1a O1a O1a O1a O1a O1a
O-M101 26 VII 1U 32 Eu16 H9 H O1a O1a1 O1a1a O1a1a O1a1 O1a1 O1a1a O1a1a O1a1a O1a1a O1a1a
O-M50 26 VII 1U 32 Eu16 H10 H O1b O1a2 O1a2 O1a2 O1a2 O1a2 O1a2 O1a2 O1a2 O1a2 O1a2
O-P31 26 VII 1U 33 Eu16 H5 I O2* O2 O2 O2 O2 O2 O2 O2 O2 O2 O2
O-M95 26 VII 1U 34 Eu16 H11 G O2a* O2a O2a O2a O2a O2a O2a O2a O2a O2a1 O2a1
O-M88 26 VII 1U 34 Eu16 H12 G O2a1 O2a1 O2a1 O2a1 O2a1 O2a1 O2a1 O2a1 O2a1 O2a1a O2a1a
O-SRY465 20 VII 1U 35 Eu16 H5 I O2b* O2b O2b O2b O2b O2b O2b O2b O2b O2b O2b
O-47z 5 VII 1U 26 Eu16 H5 I O2b1 O2b1a O2b1 O2b1 O2b1a O2b1a O2b1 O2b1 O2b1 O2b1 O2b1
O-M122 26 VII 1U 29 Eu16 H6 L O3* O3 O3 O3 O3 O3 O3 O3 O3 O3 O3
O-M121 26 VII 1U 29 Eu16 H6 L O3a O3a O3a1 O3a1 O3a1 O3a1 O3a1 O3a1 O3a1 O3a1a O3a1a
O-M164 26 VII 1U 29 Eu16 H6 L O3b O3b O3a2 O3a2 O3a2 O3a2 O3a2 O3a2 O3a2 O3a1b O3a1b
O-M159 13 VII 1U 31 Eu16 H6 L O3c O3c O3a3a O3a3a O3a3 O3a3 O3a3a O3a3a O3a3a O3a3a O3a3a
O-M7 26 VII 1U 29 Eu16 H7 L O3d* O3c O3a3b O3a3b O3a4 O3a4 O3a3b O3a3b O3a3b O3a2b O3a2b
O-M113 26 VII 1U 29 Eu16 H7 L O3d1 O3c1 O3a3b1 O3a3b1 - O3a4a O3a3b1 O3a3b1 O3a3b1 O3a2b1 O3a2b1
O-M134 26 VII 1U 30 Eu16 H8 L O3e* O3d O3a3c O3a3c O3a5 O3a5 O3a3c O3a3c O3a3c O3a2c1 O3a2c1
O-M117 26 VII 1U 30 Eu16 H8 L O3e1* O3d1 O3a3c1 O3a3c1 O3a5a O3a5a O3a3c1 O3a3c1 O3a3c1 O3a2c1a O3a2c1a
O-M162 26 VII 1U 30 Eu16 H8 L O3e1a O3d1a O3a3c1a O3a3c1a O3a5a1 O3a5a1 O3a3c1a O3a3c1a O3a3c1a O3a2c1a1 O3a2c1a1

Original Research Publications

The following research teams per their publications were represented in the creation of the YCC Tree.

Phylogenetic trees

ISOGG 2017 tree (ver. 12.244).[86]

  • O (M175)
    • O1 (F265/M1354, CTS2866, F75/M1297, F429/M1415, F465/M1422)
      • O1a (M119)
        • O1a1 (B384/Z23193)
          • O1a1a (M307.1/P203.1)
            • O1a1a1 (F446)
              • O1a1a1a (F140)
                • O1a1a1a1 (F78)
                  • O1a1a1a1a (F81)
                    • O1a1a1a1a1 (CTS2458)
                      • O1a1a1a1a1a (F533)
                        • O1a1a1a1a1a1 (F492)
                          • O1a1a1a1a1a1a (F656)
                            • O1a1a1a1a1a1a1 (A12440)
                              • O1a1a1a1a1a1a1a (A12439)
                            • O1a1a1a1a1a1a2 (A14788)
                            • O1a1a1a1a1a1a3 (F65)
                            • O1a1a1a1a1a1a4 (MF1068)
                            • O1a1a1a1a1a1a5 (Z23482)
                          • O1a1a1a1a1a1b (FGC66168)
                            • O1a1a1a1a1a1b1 (CTS11553)
                          • O1a1a1a1a1a1c (Y31266)
                            • O1a1a1a1a1a1c1 (Y31261)
                          • O1a1a1a1a1a1d (A12441)
                          • O1a1a1a1a1a1e (MF1071)
                            • O1a1a1a1a1a1e1 (MF1074)
                        • O1a1a1a1a1a2 (CTS4585)
                    • O1a1a1a1a2 (MF1075)
                • O1a1a1a2 (YP4610/Z39229)
                  • O1a1a1a2a (AM00330/AMM480/B386)
                    • O1a1a1a2a1 (AM00333/AMM483/B387)
                      • O1a1a1a2a1a (B388)
                  • O1a1a1a2b (SK1555)
              • O1a1a1b (SK1568/Z23420)
                • O1a1a1b1 (M101)
                • O1a1a1b2 (Z23392)
                  • O1a1a1b2a (Z23442)
                    • O1a1a1b2a1 (SK1571)
            • O1a1a2 (CTS52)
              • O1a1a2a (CTS701)
                • O1a1a2a1 (K644/Z23266)
          • O1a1b (CTS5726)
        • O1a2 (M110)
          • O1a2a (F3288)
            • O1a2a1 (B392)
              • O1a2a1a (B393)
        • O1a3 (Page109)
      • O1b (M268)
        • O1b1 (F2320)
          • O1b1a (M1470)
            • O1b1a1 (PK4)
              • O1b1a1a (M95)
                • O1b1a1a1 (F1803/M1348)
                  • O1b1a1a1a (F1252)
                    • O1b1a1a1a1 (F2924)
                      • O1b1a1a1a1a (M111)
                        • O1b1a1a1a1a1 (F2758)
                          • O1b1a1a1a1a1a (Z24083)
                            • O1b1a1a1a1a1a1 (Z24089)
                              • O1b1a1a1a1a1a1a (F923)
                                • O1b1a1a1a1a1a1a1 (CTS2022)
                                  • O1b1a1a1a1a1a1a1a (F1399)
                                    • O1b1a1a1a1a1a1a1a1 (F2415)
                                • O1b1a1a1a1a1a1a2 (Z24131)
                                • O1b1a1a1a1a1a1a3 (Z24100)
                              • O1b1a1a1a1a1a1b (SK1627/Z24091)
                                • O1b1a1a1a1a1a1b1 (Z39410)
                            • O1b1a1a1a1a1a2 (Z24088)
                        • O1b1a1a1a1a2 (F2890)
                          • O1b1a1a1a1a2a (Z24048)
                            • O1b1a1a1a1a2a1 (Z24050)
                          • O1b1a1a1a1a2b (Z24014)
                      • O1b1a1a1a1b (CTS5854)
                        • O1b1a1a1a1b1 (Z23810)
                          • O1b1a1a1a1b1a (CTS7399)
                            • O1b1a1a1a1b1a1 (FGC19713/Y14026)
                              • O1b1a1a1a1b1a1a (Z23849)
                                • O1b1a1a1a1b1a1a1 (FGC61038)
                          • O1b1a1a1a1b1b (CTS651)
                            • O1b1a1a1a1b1b1 (CTS9884)
                        • O1b1a1a1a1b2 (F4229)
                          • O1b1a1a1a1b2a (F809)
                            • O1b1a1a1a1b2a1 (F2517)
                    • O1b1a1a1a2 (SK1630)
                      • O1b1a1a1a2a (SK1636)
                  • O1b1a1a1b (F789/M1283)
                    • O1b1a1a1b1 (FGC29900/Y9322/Z23667)
                      • O1b1a1a1b1a (B426/FGC29896/Y9033/Z23671)
                        • O1b1a1a1b1a1 (FGC29907/YP3930)
                        • O1b1a1a1b1a2 (B427/Z23680)
                      • O1b1a1a1b1b (Z39485)
                      • O1b1a1a1b1c (B418)
                    • O1b1a1a1b2 (SK1646)
                • O1b1a1a2 (CTS350)
                • O1b1a1a3 (Page103)
              • O1b1a1b (F838)
                • O1b1a1b1 (F1199)
            • O1b1a2 (Page59)
              • O1b1a2a (F993)
                • O1b1a2a1 (F1759)
                  • O1b1a2a1a (CTS1127)
              • O1b1a2b (F417/M1654)
                • O1b1a2b1 (F840)
                  • O1b1a2b1a (F1127)
                • O1b1a2b2 (CTS1451)
              • O1b1a2c (CTS9996)
        • O1b2 (P49, M176)
          • O1b2a (F1942/Page92)
            • O1b2a1 (CTS9259)
              • O1b2a1a (F1204)
                • O1b2a1a1 (CTS713)
                  • O1b2a1a1a (CTS1875)
                    • O1b2a1a1a1 (CTS10682)
                  • O1b2a1a1b (Z24598)
                  • O1b2a1a1c (CTS203)
                • O1b2a1a2 (F2868)
                  • O1b2a1a2a (L682)
                    • O1b2a1a2a1 (CTS723)
                      • O1b2a1a2a1a (CTS7620)
                      • O1b2a1a2a1b (A12446)
                        • O1b2a1a2a1b1 (PH40)
                  • O1b2a1a2b (F940)
                • O1b2a1a3 (CTS10687)
                  • O1b2a1a3a (CTS1215)
              • O1b2a1b (CTS562)
            • O1b2a2 (Page90)
    • O2 (M122)
      • O2a (M324)
        • O2a1 (L127.1)
          • O2a1a (F1876/Page127)
            • O2a1a1 (F2159)
              • O2a1a1a (F1867/Page124)
                • O2a1a1a1 (F852)
                  • O2a1a1a1a (F2266)
                    • O2a1a1a1a1 (L599)
                      • O2a1a1a1a1a (Z43961)
                        • O2a1a1a1a1a1 (Z43963)
                  • O2a1a1a1b (F854)
                    • O2a1a1a1b1 (Z43966)
                  • O2a1a1a1c (Page130)
              • O2a1a1b (F915)
                • O2a1a1b1 (F1478)
                  • O2a1a1b1a (PF5390)
                    • O2a1a1b1a1 (CTS1936)
                    • O2a1a1b1a1a (Z43975)
                    • O2a1a1b1a2 (FGC33994)
          • O2a1b (M164)
          • O2a1c (IMS-JST002611)
            • O2a1c1 (F18)
              • O2a1c1a (F117)
                • O2a1c1a1 (F13)
                  • O2a1c1a1a (F11)
                    • O2a1c1a1a1 (F632)
                      • O2a1c1a1a1a (F110/M11115)
                        • O2a1c1a1a1a1 (F17)
                          • O2a1c1a1a1a1a (F377)
                            • O2a1c1a1a1a1a1 (F1095)
                              • O2a1c1a1a1a1a1a (F856)
                                • O2a1c1a1a1a1a1a1 (F1418)
                                • O2a1c1a1a1a1a1a2 (Z25097)
                            • O2a1c1a1a1a1a2 (CTS7501)
                          • O2a1c1a1a1a1b (F793)
                        • O2a1c1a1a1a2 (Y20951)
                          • O2a1c1a1a1a2a (Y20932)
                    • O2a1c1a1a2 (F38)
                    • O2a1c1a1a3 (F12)
                    • O2a1c1a1a4 (F930)
                      • O2a1c1a1a4a (F2685)
                    • O2a1c1a1a5 (F1365/M5420/PF1558)
                      • O2a1c1a1a5a (Y15976)
                        • O2a1c1a1a5a1 (Y16154)
                          • O2a1c1a1a5a1a (Y26383)
                            • O2a1c1a1a5a1a1 (SK1686)
                      • O2a1c1a1a5b (FGC54486)
                        • O2a1c1a1a5b1 (FGC54507)
                    • O2a1c1a1a6 (CTS12877)
                      • O2a1c1a1a6a (F2527)
                        • O2a1c1a1a6a1 (CTS5409)
                        • O2a1c1a1a6a2 (F2941)
                    • O2a1c1a1a7 (F723)
                    • O2a1c1a1a8 (CTS2107)
                    • O2a1c1a1a9 (SK1691)
                  • O2a1c1a1b (PH203)
              • O2a1c1b (F449)
                • O2a1c1b1 (F238)
                  • O2a1c1b1a (F134)
                    • O2a1c1b1a1 (F1273)
                    • O2a1c1b1a2 (F724)
                • O2a1c1b2 (F1266)
              • O2a1c1c (CTS498)
            • O2a1c2 (FGC3750/SK1673)
        • O2a2 (IMS-JST021354/P201)
          • O2a2a (M188)
            • O2a2a1 (F2588)
              • O2a2a1a (CTS445)
                • O2a2a1a1 (CTS201)
                  • O2a2a1a1a (M159/Page96)
                • O2a2a1a2 (M7)
                  • O2a2a1a2a (F1276)
                    • O2a2a1a2a1 (CTS6489)
                      • O2a2a1a2a1a (F1275)
                        • O2a2a1a2a1a1 (M113)
                        • O2a2a1a2a1a2 (N5)
                        • O2a2a1a2a1a3 (Z25400)
                    • O2a2a1a2a2 (F1863)
                      • O2a2a1a2a2a (F1134)
                        • O2a2a1a2a2a1 (F1262)
                  • O2a2a1a2b (Y26403)
              • O2a2a1b (F1837)
            • O2a2a2 (F879)
              • O2a2a2a (F1226)
                • O2a2a2a1 (F2859)
          • O2a2b (P164)
            • O2a2b1 (M134)
              • O2a2b1a (F450/M1667)
                • O2a2b1a1 (M117/Page23)
                  • O2a2b1a1a (M133)
                    • O2a2b1a1a1 (F438)
                      • O2a2b1a1a1a (Y17728)
                        • O2a2b1a1a1a1 (F155)
                          • O2a2b1a1a1a1a (F813/M6539)
                            • O2a2b1a1a1a1a1 (Y20928)
                        • O2a2b1a1a1a2 (F1754)
                          • O2a2b1a1a1a2a (F2137)
                            • O2a2b1a1a1a2a1 (F1442)
                              • O2a2b1a1a1a2a1a (F1123)
                                • O2a2b1a1a1a2a1a1 (F1369)
                            • O2a2b1a1a1a2a2 (A16636)
                        • O2a2b1a1a1a3 (Z25907)
                    • O2a2b1a1a2 (FGC23469/Z25852)
                      • O2a2b1a1a2a (F310)
                        • O2a2b1a1a2a1 (F402)
                          • O2a2b1a1a2a1a (F1531)
                    • O2a2b1a1a3 (CTS7634)
                      • O2a2b1a1a3a (F317)
                        • O2a2b1a1a3a1 (F3039)
                        • O2a2b1a1a3a2 (Y29861)
                      • O2a2b1a1a3b (CTS5488)
                    • O2a2b1a1a4 (Z25853)
                      • O2a2b1a1a4a (CTS5492)
                        • O2a2b1a1a4a1 (CTS6987)
                          • O2a2b1a1a4a1a (Z42620)
                        • O2a2b1a1a4a2 ( F20963)
                    • O2a2b1a1a5 (CTS10738/M1707)
                      • O2a2b1a1a5a (CTS9678)
                        • O2a2b1a1a5a1 (Z39663)
                        • O2a2b1a1a5a2 (M1513)
                      • O2a2b1a1a5b (A9457)
                        • O2a2b1a1a5b1 (F17158)
                    • O2a2b1a1a6 (CTS4658)
                      • O2a2b1a1a6a (CTS5308)
                      • O2a2b1a1a6b (Z25928)
                        • O2a2b1a1a6b1 (SK1730/Z25982)
                          • O2a2b1a1a6b1a (Z26030)
                          • O2a2b1a1a6b1b (Z26010)
                        • O2a2b1a1a6b2 (A9462)
                        • O2a2b1a1a6b3 (B456)
                    • O2a2b1a1a7 (YP4864)
                      • O2a2b1a1a7a (Z44068)
                        • O2a2b1a1a7a1 (F5525/SK1748)
                      • O2a2b1a1a7b (Z44071)
                    • O2a2b1a1a8 (Z44091)
                      • O2a2b1a1a8a (Z44092)
                  • O2a2b1a1b (CTS4960)
                • O2a2b1a2 (F114)
                  • O2a2b1a2a (F79)
                    • O2a2b1a2a1 (F46/Y15)
                      • O2a2b1a2a1a (FGC16847/Z26091)
                        • O2a2b1a2a1a1 (F48)
                          • O2a2b1a2a1a1a (F152)
                            • O2a2b1a2a1a1a1 (F2505)
                          • O2a2b1a2a1a1b (CTS3149)
                        • O2a2b1a2a1a2 (F242)
                          • O2a2b1a2a1a2a (CTS4266)
                            • O2a2b1a2a1a2a1 (Z26108)
                              • O2a2b1a2a1a2a1a (F2173)
                        • O2a2b1a2a1a3 (F2887)
                          • O2a2b1a2a1a3a (F3607)
                            • O2a2b1a2a1a3a1 (F3525)
                          • O2a2b1a2a1a3b (CTS3763)
                            • O2a2b1a2a1a3b1 (A9472)
                            • O2a2b1a2a1a3b2 (FGC16863/Y7110)
                              • O2a2b1a2a1a3b2a (L1360)
                                • O2a2b1a2a1a3b2a1 (FGC16889)
                              • O2a2b1a2a1a3b2b (SK1768/Y7112/Z26257)
                                • O2a2b1a2a1a3b2b1 (F4249)
                                  • O2a2b1a2a1a3b2b1a (FGC23868)
                                • O2a2b1a2a1a3b2b2 (CTS335)
                      • O2a2b1a2a1b (CTS53)
                        • O2a2b1a2a1b1 (CTS6373)
                          • O2a2b1a2a1b1a (A9473)
                      • O2a2b1a2a1c (F3386)
                      • O2a2b1a2a1d (Y29828)
                        • O2a2b1a2a1d1 (F735)
                          • O2a2b1a2a1d1a (FGC34973)
                          • O2a2b1a2a1d1b (F1739)
                  • O2a2b1a2b (F743)
                    • O2a2b1a2b1 (CTS8481)
                      • O2a2b1a2b1a (CTS4325)
                        • O2a2b1a2b1a1 (A16629)
                        • O2a2b1a2b1a2 (CTS682)
                    • O2a2b1a2b2 (F748)
                      • O2a2b1a2b2a (F728)
                  • O2a2b1a2c (Page101)
            • O2a2b2 (AM01822/F3223)
              • O2a2b2a (AM01856/F871)
                • O2a2b2a1 (N7)
                  • O2a2b2a1a (F4110)
                    • O2a2b2a1a1 (F4068)
                    • O2a2b2a1a2 (SK1780)
                  • O2a2b2a1b (F4124)
                    • O2a2b2a1b1 (IMS-JST008425p6)
                    • O2a2b2a1b2 (BY15188)
                      • O2a2b2a1b2a (F16411)
                • O2a2b2a2 (AM01845/F706)
                  • O2a2b2a2a (F717)
                    • O2a2b2a2a1 (F3612)
                    • O2a2b2a2a2 (SK1783)
                  • O2a2b2a2b (AM01847/B451)
                    • O2a2b2a2b1 (A17418)
                    • O2a2b2a2b2 (AM01756)
                      • O2a2b2a2b2a (B450)
                      • O2a2b2a2b2b (AM00472/B452)
                        • O2a2b2a2b2b1 (F18942)
                      • O2a2b2a2b2c (A16427)
              • O2a2b2b (A16433)
                • O2a2b2b1 (A16438)
                  • O2a2b2b1a (SK1775)
                    • O2a2b2b1a1 (SK1774)
                  • O2a2b2b1b (A16440)
        • O2a3 (M300)
        • O2a4 (M333)
      • O2b (F742)
        • O2b1 (F1150)
          • O2b1a (F837)
            • O2b1a1 (F1025)
        • O2b2 (F1055)
          • O2b2a (F3021)

See also

Genetics

Y-DNA O subclades

Y-DNA backbone tree

Notes

  1. O-M175(xM119,M95,M122) is sometimes incorrectly called "O*".
  2. The outlier Kadai branch is called "Kra" by Thai linguist Weera Ostapirat and "Geyang" by Chinese linguists.

References

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  41. 41.000 41.001 41.002 41.003 41.004 41.005 41.006 41.007 41.008 41.009 41.010 41.011 41.012 41.013 41.014 41.015 41.016 41.017 41.018 41.019 41.020 41.021 41.022 41.023 41.024 41.025 41.026 41.027 41.028 41.029 41.030 41.031 41.032 41.033 41.034 41.035 41.036 41.037 41.038 41.039 41.040 41.041 41.042 41.043 41.044 41.045 41.046 41.047 41.048 41.049 41.050 41.051 41.052 41.053 41.054 41.055 41.056 41.057 41.058 41.059 41.060 41.061 41.062 41.063 41.064 41.065 41.066 41.067 41.068 41.069 41.070 41.071 41.072 41.073 41.074 41.075 41.076 41.077 41.078 41.079 41.080 41.081 41.082 41.083 41.084 41.085 41.086 41.087 41.088 41.089 41.090 41.091 41.092 41.093 41.094 41.095 41.096 41.097 41.098 41.099 41.100 41.101 41.102 41.103 41.104 41.105 41.106 41.107 41.108 41.109 41.110 41.111 41.112 41.113 41.114 41.115 41.116 41.117 41.118 41.119 41.120 41.121 41.122 41.123 41.124 41.125 41.126 41.127 41.128 41.129 41.130 41.131 41.132 41.133 41.134 41.135 41.136 41.137 41.138 41.139 41.140 41.141 41.142 41.143 41.144 41.145 41.146 41.147 41.148 41.149 41.150 41.151 41.152 41.153 41.154 41.155 41.156 41.157 41.158 41.159 41.160 41.161 41.162 41.163 41.164 41.165 41.166 41.167 41.168 41.169 41.170 41.171 41.172 41.173 41.174 41.175 41.176 41.177 41.178 41.179 41.180 41.181 41.182 41.183 41.184 41.185 41.186 41.187 41.188 41.189 41.190 41.191 41.192 41.193 41.194 41.195 41.196 41.197 41.198 41.199 41.200 41.201 41.202 41.203 41.204 41.205 41.206 41.207 41.208 41.209 41.210 41.211 41.212 41.213 41.214 41.215 41.216 41.217 41.218 41.219 41.220 41.221 41.222 41.223 41.224 41.225 41.226 41.227 41.228 41.229 41.230 41.231 41.232 41.233 41.234 41.235 41.236 41.237 41.238 41.239 41.240 41.241 41.242 41.243 41.244 41.245 41.246 41.247 41.248 41.249 41.250 41.251 41.252 41.253 41.254 41.255 41.256 41.257 41.258 41.259 41.260 41.261 41.262 41.263 41.264 41.265 41.266 41.267 41.268 41.269 41.270 41.271 41.272 41.273 41.274 41.275 41.276 41.277 41.278 41.279 41.280 41.281 41.282 41.283 41.284 41.285 41.286 41.287 41.288 41.289 41.290 41.291 41.292 41.293 41.294 41.295 41.296 41.297 41.298 41.299 41.300 41.301 41.302 41.303 41.304 41.305 41.306 41.307 41.308 41.309 41.310 41.311 41.312 41.313 41.314 41.315 41.316 41.317 41.318 41.319 41.320 41.321 41.322 41.323 41.324 41.325 41.326 41.327 41.328 41.329 41.330 41.331 41.332 41.333 41.334 41.335 41.336 41.337 41.338 41.339 41.340 41.341 41.342 41.343 41.344 41.345 41.346 41.347 41.348 41.349 41.350 41.351 41.352 41.353 41.354 41.355 41.356 41.357 41.358 41.359 41.360 41.361 41.362 41.363 41.364 41.365 41.366 41.367 41.368 41.369 41.370 41.371 41.372 41.373 41.374 41.375 41.376 41.377 41.378 41.379 41.380 41.381 41.382 41.383 41.384 41.385 41.386 41.387 41.388 41.389 41.390 41.391 41.392 41.393 41.394 41.395 41.396 41.397 41.398 41.399 41.400 41.401 41.402 41.403 41.404 41.405 41.406 41.407 41.408 41.409 41.410 41.411 41.412 41.413 41.414 41.415 41.416 41.417 41.418 41.419 41.420 41.421 41.422 41.423 41.424 41.425 41.426 41.427 41.428 41.429 41.430 41.431 41.432 41.433 41.434 41.435 41.436 41.437 41.438 41.439 41.440 41.441 41.442 41.443 41.444 41.445 41.446 41.447 41.448 41.449 41.450 41.451 41.452 YFull Haplogroup YTree v5.04 at 16 May 2017
  42. 42.000 42.001 42.002 42.003 42.004 42.005 42.006 42.007 42.008 42.009 42.010 42.011 42.012 42.013 42.014 42.015 42.016 42.017 42.018 42.019 42.020 42.021 42.022 42.023 42.024 42.025 42.026 42.027 42.028 42.029 42.030 42.031 42.032 42.033 42.034 42.035 42.036 42.037 42.038 42.039 42.040 42.041 42.042 42.043 42.044 42.045 42.046 42.047 42.048 42.049 42.050 42.051 42.052 42.053 42.054 42.055 42.056 42.057 42.058 42.059 42.060 42.061 42.062 42.063 42.064 42.065 42.066 42.067 42.068 42.069 42.070 42.071 42.072 42.073 42.074 42.075 42.076 42.077 42.078 42.079 42.080 42.081 42.082 42.083 42.084 42.085 42.086 42.087 42.088 42.089 42.090 42.091 42.092 42.093 42.094 42.095 42.096 42.097 42.098 42.099 42.100 42.101 42.102 42.103 42.104 42.105 42.106 42.107 42.108 42.109 42.110 42.111 42.112 42.113 42.114 42.115 42.116 42.117 42.118 42.119 42.120 42.121 42.122 42.123 42.124 42.125 42.126 42.127 42.128 42.129 42.130 42.131 42.132 42.133 42.134 42.135 42.136 42.137 42.138 42.139 42.140 42.141 42.142 42.143 42.144 42.145 42.146 Phylogenetic tree of Haplogroup O at 23mofang
  43. 43.00 43.01 43.02 43.03 43.04 43.05 43.06 43.07 43.08 43.09 43.10 43.11 43.12 43.13 43.14 43.15 43.16 43.17 43.18 43.19 43.20 43.21 43.22 43.23 43.24 43.25 43.26 43.27 43.28 43.29 43.30 43.31 43.32 43.33 43.34 43.35 43.36 43.37 43.38 43.39 43.40 43.41 Phylogenetic tree of Haplogroup O-F175 at TheYtree
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  45. 45.00 45.01 45.02 45.03 45.04 45.05 45.06 45.07 45.08 45.09 45.10 45.11 45.12 45.13 45.14 45.15 45.16 45.17 45.18 45.19 45.20 45.21 45.22 45.23 45.24 45.25 45.26 45.27 45.28 45.29 45.30 45.31 45.32 45.33 45.34 45.35 45.36 45.37 45.38 45.39 45.40 45.41 Macholdt, Enrico; Arias, Leonardo; Duong, Nguyen Thuy; Ton, Nguyen Dang; Van Phong, Nguyen; Schröder, Roland; Pakendorf, Brigitte; Van Hai, Nong; Stoneking, Mark (May 2020). "The paternal and maternal genetic history of Vietnamese populations". European Journal of Human Genetics. 28 (5): 636–645. doi:10.1038/s41431-019-0557-4. PMC 7171127. PMID 31827276.
  46. 46.0 46.1 O Y-Haplogroup Project at Family Tree DNA
  47. 47.00 47.01 47.02 47.03 47.04 47.05 47.06 47.07 47.08 47.09 47.10 47.11 47.12 47.13 47.14 47.15 47.16 47.17 47.18 47.19 47.20 47.21 47.22 47.23 47.24 47.25 47.26 47.27 47.28 47.29 47.30 47.31 Y-DNA Haplotree at Family Tree DNA
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  49. 49.0 49.1 49.2 49.3 49.4 49.5 49.6 49.7 49.8 Zhang, Xiaoming; Kampuansai, Jatupol; Qi, Xuebin; Yan, Shi; Yang, Zhaohui; Serey, Bun; Sovannary, Tuot; Bunnath, Long; Aun, Hong Seang; Samnom, Ham; Kutanan, Wibhu; Luo, Xin; Liao, Shiyu; Kangwanpong, Daoroong; Jin, Li; Shi, Hong; Su, Bing (27 June 2014). "An Updated Phylogeny of the Human Y-Chromosome Lineage O2a-M95 with Novel SNPs". PLOS ONE. 9 (6): e101020. Bibcode:2014PLoSO...9j1020Z. doi:10.1371/journal.pone.0101020. PMC 4074153. PMID 24972021.
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  51. 51.000 51.001 51.002 51.003 51.004 51.005 51.006 51.007 51.008 51.009 51.010 51.011 51.012 51.013 51.014 51.015 51.016 51.017 51.018 51.019 51.020 51.021 51.022 51.023 51.024 51.025 51.026 51.027 51.028 51.029 51.030 51.031 51.032 51.033 51.034 51.035 51.036 51.037 51.038 51.039 51.040 51.041 51.042 51.043 51.044 51.045 51.046 51.047 51.048 51.049 51.050 51.051 51.052 51.053 51.054 51.055 51.056 51.057 51.058 51.059 51.060 51.061 51.062 51.063 51.064 51.065 51.066 51.067 51.068 51.069 51.070 51.071 51.072 51.073 51.074 51.075 51.076 51.077 51.078 51.079 51.080 51.081 51.082 51.083 51.084 51.085 51.086 51.087 51.088 51.089 51.090 51.091 51.092 51.093 51.094 51.095 51.096 51.097 51.098 51.099 51.100 51.101 Kutanan, Wibhu; Kampuansai, Jatupol; Srikummool, Metawee; Brunelli, Andrea; Ghirotto, Silvia; Arias, Leonardo; Macholdt, Enrico; Hübner, Alexander; Schröder, Roland; Stoneking, Mark (1 July 2019). "Contrasting Paternal and Maternal Genetic Histories of Thai and Lao Populations". Molecular Biology and Evolution. 36 (7): 1490–1506. doi:10.1093/molbev/msz083. PMC 6573475. PMID 30980085.
  52. Xia, Zi-Yang; Yan, Shi; Wang, Chuan-Chao; Zheng, Hong-Xiang; Zhang, Fan; Liu, Yu-Chi; Yu, Ge; Yu, Bin-Xia; Shu, Li-Li; Jin, Li (9 August 2019). "Inland-coastal bifurcation of southern East Asians revealed by Hmong-Mien genomic history". bioRxiv 10.1101/730903.S2CID 202028061
  53. 53.0 53.1 53.2 53.3 53.4 53.5 53.6 53.7 Karafet, T. M.; Hallmark, B.; Cox, M. P.; Sudoyo, H.; Downey, S.; Lansing, J. S.; Hammer, M. F. (1 August 2010). "Major East-West Division Underlies Y Chromosome Stratification across Indonesia". Molecular Biology and Evolution. 27 (8): 1833–1844. doi:10.1093/molbev/msq063. PMID 20207712.
  54. 54.0 54.1 54.2 54.3 54.4 54.5 54.6 54.7 54.8 Trejaut, Jean A; Poloni, Estella S; Yen, Ju-Chen; Lai, Ying-Hui; Loo, Jun-Hun; Lee, Chien-Liang; He, Chun-Lin; Lin, Marie (2014). "Taiwan Y-chromosomal DNA variation and its relationship with Island Southeast Asia". BMC Genetics. 15 (1): 77. doi:10.1186/1471-2156-15-77. PMC 4083334. PMID 24965575.
  55. 55.0 55.1 55.2 55.3 55.4 55.5 55.6 Xue, Yali; Zerjal, Tatiana; Bao, Weidong; Zhu, Suling; Shu, Qunfang; Xu, Jiujin; Du, Ruofu; Fu, Songbin; Li, Pu; Hurles, Matthew E; Yang, Huanming; Tyler-Smith, Chris (1 April 2006). "Male Demography in East Asia: A North–South Contrast in Human Population Expansion Times". Genetics. 172 (4): 2431–2439. doi:10.1534/genetics.105.054270. PMC 1456369. PMID 16489223.
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  57. 57.0 57.1 57.2 57.3 57.4 57.5 Su, Bing; Xiao, Junhua; Underhill, Peter; Deka, Ranjan; Zhang, Weiling; Akey, Joshua; Huang, Wei; Shen, Di; Lu, Daru; Luo, Jingchun; Chu, Jiayou; Tan, Jiazhen; Shen, Peidong; Davis, Ron; Cavalli-Sforza, Luca; Chakraborty, Ranajit; Xiong, Momiao; Du, Ruofu; Oefner, Peter; Chen, Zhu; Jin, Li (December 1999). "Y-Chromosome Evidence for a Northward Migration of Modern Humans into Eastern Asia during the Last Ice Age". The American Journal of Human Genetics. 65 (6): 1718–1724. doi:10.1086/302680. PMC 1288383. PMID 10577926.
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  59. 59.0 59.1 59.2 He, Jun-Dong; Peng, Min-Sheng; Quang, Huy Ho; Dang, Khoa Pham; Trieu, An Vu; Wu, Shi-Fang; Jin, Jie-Qiong; Murphy, Robert W.; Yao, Yong-Gang; Zhang, Ya-Ping (7 May 2012). "Patrilineal Perspective on the Austronesian Diffusion in Mainland Southeast Asia". PLOS ONE. 7 (5): e36437. Bibcode:2012PLoSO...736437H. doi:10.1371/journal.pone.0036437. PMC 3346718. PMID 22586471.
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  61. 61.0 61.1 61.2 61.3 61.4 61.5 Cai, Xiaoyun; Qin, Zhendong; Wen, Bo; Xu, Shuhua; Wang, Yi; Lu, Yan; Wei, Lanhai; Wang, Chuanchao; Li, Shilin; Huang, Xingqiu; Jin, Li; Li, Hui (31 August 2011). "Human Migration through Bottlenecks from Southeast Asia into East Asia during Last Glacial Maximum Revealed by Y Chromosomes". PLOS ONE. 6 (8): e24282. Bibcode:2011PLoSO...624282C. doi:10.1371/journal.pone.0024282. PMC 3164178. PMID 21904623.
  62. Wen, Bo (2004). "The origin of Mosuo people as revealed by mtDNA and Y chromosome variation". Science in China Series C. 47 (1): 1–10. doi:10.1360/02yc0207. PMID 15382670. S2CID 7999778.
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  64. 64.00 64.01 64.02 64.03 64.04 64.05 64.06 64.07 64.08 64.09 64.10 64.11 64.12 64.13 64.14 64.15 64.16 64.17 64.18 64.19 64.20 64.21 64.22 64.23 64.24 64.25 64.26 64.27 Yao X, Tang S, Bian B, Wu X, Chen G, Wang CC (April 2017). "Improved phylogenetic resolution for Y-chromosome Haplogroup O2a1c-002611". Scientific Reports. 7 (1): 1146. Bibcode:2017NatSR...7.1146Y. doi:10.1038/s41598-017-01340-z. PMC 5430735. PMID 28442769.
  65. "O-M159单倍群详情".
  66. "O-Mf22947单倍群详情".
  67. "O-Z25518单倍群详情".
  68. "O-Mf14135单倍群详情".
  69. "夏代东部大族祖源分析-23魔方祖源基因检测".
  70. 70.00 70.01 70.02 70.03 70.04 70.05 70.06 70.07 70.08 70.09 70.10 70.11 70.12 70.13 70.14 70.15 70.16 70.17 70.18 70.19 70.20 70.21 70.22 70.23 70.24 70.25 70.26 70.27 70.28 70.29 70.30 70.31 70.32 Time Tree of Y-DNA haplogroup O-M175 at FamilyTreeDNA Discover
  71. Li, Hui; Huang, Ying; Mustavich, Laura F.; Zhang, Fan; Tan, Jing-Ze; Wang, Ling-E; Qian, Ji; Gao, Meng-He; Jin, Li (November 2007). "Y chromosomes of prehistoric people along the Yangtze River". Human Genetics. 122 (3–4): 383–388. doi:10.1007/s00439-007-0407-2. PMID 17657509. S2CID 2533393.
  72. "O-Mf9858单倍群详情".
  73. "O-F1262单倍群详情".
  74. "O-Mf106428单倍群详情".
  75. 75.0 75.1 75.2 75.3 75.4 75.5 Kutanan, Wibhu; Shoocongdej, Rasmi; Srikummool, Metawee; Hübner, Alexander; Suttipai, Thanatip; Srithawong, Suparat; Kampuansai, Jatupol; Stoneking, Mark (November 2020). "Cultural variation impacts paternal and maternal genetic lineages of the Hmong-Mien and Sino-Tibetan groups from Thailand". European Journal of Human Genetics. 28 (11): 1563–1579. doi:10.1038/s41431-020-0693-x. PMC 7576213. PMID 32690935.
  76. "O-Mf48275单倍群详情".
  77. "O-F14832单倍群详情".
  78. "O-Y140772单倍群详情".
  79. "O-Z25400单倍群详情".
  80. "O-Mf36985单倍群详情".
  81. "O-Mf193618单倍群详情".
  82. Huang, Xiufeng; Xia, Zi-Yang; Bin, Xiaoyun; He, Guanglin; Guo, Jianxin (30 June 2022). "Genomic Insights Into the Demographic History of the Southern Chinese". Frontiers in Ecology and Evolution. 10. Frontiers Media SA. doi:10.3389/fevo.2022.853391. ISSN 2296-701X.
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  84. Wei, Lan-Hai; Yan, Shi; Teo, Yik-Ying; Huang, Yun-Zhi; Wang, Ling-Xiang; Yu, Ge; Saw, Woei-Yuh; Ong, Rick Twee-Hee; Lu, Yan; Zhang, Chao; Xu, Shu-Hua; Jin, Li; Li, Hui (5 April 2017). "Phylogeography of Y-chromosome haplogroup O3a2b2-N6 reveals patrilineal traces of Austronesian populations on the eastern coastal regions of Asia". PLOS ONE. 12 (4): e0175080. Bibcode:2017PLoSO..1275080W. doi:10.1371/journal.pone.0175080. PMC 5381892. PMID 28380021.
  85. 85.0 85.1 85.2 85.3 Mondal, Mayukh; Casals, Ferran; Xu, Tina; Dall'Olio, Giovanni M; Pybus, Marc; Netea, Mihai G; Comas, David; Laayouni, Hafid; Li, Qibin; Majumder, Partha P; Bertranpetit, Jaume (September 2016). "Genomic analysis of Andamanese provides insights into ancient human migration into Asia and adaptation". Nature Genetics. 48 (9): 1066–1070. doi:10.1038/ng.3621. hdl:10230/34401. PMID 27455350. S2CID 205352099.
  86. "Y-DNA Haplogroup D-M174 and its Subclades - 2017".

Sources for conversion tables

Further reading

External links